Marsh Muhly
Muhlenbergia racemosa
Grass family (Poaceae)

Spike-like PanicleDescription: This native perennial grass is 1-3' long and erect to sprawling. The branching culms are light green, terete, and glabrous. Abundant alternate leaves occur along the length of each culm; lower leaves tend to wither away. The leaf blades are up to 4" long and 5 mm. across; they are dull grey-green and hairless. The blades are ascending to widely spreading and either floppy or somewhat stiff. The leaf sheaths are dull grey green, hairless, somewhat flattened (or keeled), and often coming loose from the culms. At the junction of each blade and sheath, the ligule consists of a papery truncate membrane about 1 mm. in length. The small nodes are swollen and hairless. Each upper culm terminates in a narrow panicle of spikelets up to 5" long and ½" across. Initially, this panicle has a spike-like appearance, but it becomes a little more broad with age as its secondary branches become less appressed against the central axis. These secondary branches are up to 1½" long. The spikelets are densely distributed along the secondary branches; they are light green or whitish green, becoming light brown with age. Each spikelet is 6-8 mm. long, consisting of a pair of awned glumes and a single lemma. The glumes are 6-8 mm. long (including their awns), keeled-linear in shape, and mostly membranous, except for their central veins and awns. The lemma is 3-4 mm. long, linear-lanceolate in shape, and pale green; there is a tuft of fine hairs at its base. The tiny anthers of each wind-pollinated floret are dull white and about 0.5 mm. long. The blooming period occurs during the late summer and the grains of the spikelets ripen during the fall. Disarticulation is above the glumes. Each narrow grain is 1.5–2.0 mm. long. The root system is fibrous, long-rhizomatous, and rather shallow. Loose colonies of plants are often formed from the rhizomes.

Cultivation: Generally, this grass prefers full or partial sun and a moist soil containing loam, sandy loam, or silt. However, some local ecotypes of this species are adapted to drier sites; the latter tend to have a more erect habit and stiffer leaf blades.

Range & Habitat: Marsh Muhly is occasional in the northern 3 tiers of counties and the western half of Illinois, but it is uncommon or absent elsewhere in the state (see Distribution Map). Illinois lies toward the eastern range-limit of this species. Habitats are highly variable; they include moist to dry-mesic prairiesCulms & Leaves (particularly in disturbed areas), thin woodlands and savannas, low areas along rivers, roadside ditches, and edges of marshes. Usually, these habitats are somewhat degraded from a history of disturbance.

Faunal Associations: The leafhoppers Flexamia imputans and Plesiommata tripunctata feed on Muhlenbergia spp. in open woodlands and other sheltered situations; sometimes Marsh Muhly occurs in these kinds of habitats. The foliage is edible to cattle and other livestock while it is young. Because this grass can form rather dense colonies, it can provide significant cover for small mammals and other wildlife.

Photographic Location: The wildflower garden of the webmaster in Urbana, Illinois.

Comments: Marsh Muhly is a rather ordinary-looking grass that resembles Muhlenbergia frondosa (Common Satin Grass) and several other species in this genus. It can be distinguished from them primarily by its awned glumes, which are 6-8 mm. long. Other Muhlenbergia spp. in Illinois have glumes that lack awns, although their lemmas are sometimes awned. An exception is Muhlenbergia glomerata (Wild Timothy), which has awned glumes of the same length as Marsh Muhly; these two species are very difficult to distinguish. However, the anthers of Marsh Muhly are much smaller (about 0.5 mm. in length) than those of Wild Timothy (about 1.0–1.5 mm. in length), while the papery ligules of Marsh Muhly are longer (about 1.0 mm. in length) than those of Wild Timothy (about 0.5 mm. in length or less). There have been some attempts to make generalizations about the differing habitats that these two species are supposed to prefer, but this is rather risky because fidelity to any particular habitat appears to be low.

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