Description:
This is a woody vine with trailing stems up to 8' long that branch
occasionally. The stems are light green to red, angular or terete, and
sparsely to moderately covered with bristly hairs. In addition,
sometimes softer hairs and widely scattered small prickles
are present along the stems. Alternate trifoliate leaves occur
along these stems on long petioles up to 4" long. On rare occasions,
some leaves may have 5 leaflets, instead of the usual 3 leaflets.
Individual leaflets are 1-2½" long and about one-half as much across;
they are ovate to obovate and coarsely toothed along their middle to
upper margins. The upper leaflet surface is medium green, hairless, and
somewhat shiny, while the lower surface is more pale and sometimes
softly hairy. The terminal leaflet of each trifoliate leaf has a short
petiolule (basal stalklet) about 1/8" long, while the lateral leaflets
are sessile. Occasionally, small cymes of 3-6 flowers are produced from
axils of the leaves on peduncles up to 6" long. The peduncles are light
green to reddish green and sparsely to moderately covered with small
bristles or hairs. Each flower is ½-¾" across, consisting of 5 white
petals, 5 light green sepals that are united at the base, a ring of
numerous stamens, and a compound pistil at its center that is
light green. The petals are oblong-elliptic in
shape, while the sepals are ovate and softly hairy. The
blooming
period occurs during early to mid-summer for about 3 weeks. Fertile
flowers are replaced by compound drupes that are ¼-½" in length at
maturity. Immature drupes are light green or white, becoming red during
an intermediate stage, and finally dark purple or black when they are
ripe. Each drupe consists of a cluster of small drupelets; each fleshy
druplet contains a single seed. Usually, the flavor of mature
drupes is sour. The root system is woody and branching. Sometimes the
tips of trailing stems develop rootlets on moist ground, enabling this
vine to spread vegetatively. The leaves are semi-evergreen and often
become reddish during the fall or winter.
Cultivation:
The preference is dappled sunlight, wet to moist
conditions, and an acidic soil containing sand or peat.
Range
& Habitat: The native Swamp Dewberry is occasional
in northeast
Illinois and either rare or absent elsewhere in the state. Habitats
include swamps, moist sandy woodlands, moist sandy
thickets, moist sand prairies and sandy shrub prairies, edges
of marshes, and bogs. This
vine is found in both degraded and higher quality habitats,
favoring areas that have been burned over by fire during
periods of drought.
Faunal
Associations: The flowers of Swamp Dewberry are visited
primarily by
bees, including both long-tongued and short-tongued bees. Other insects
that may visit the flowers include Syrphid flies, bee flies, small
butterflies, and skippers. Both nectar and pollen are available as
floral rewards to such visitors. The remaining information about
floral-faunal relationship applies primarily to
Rubus spp. in
general,
which consist of blackberries, raspberries, and dewberries. Many
insects feed on the leaves, stems, fruit, plant juices, and other parts
of these woody plants. These species include leaf beetles, aphids,
leafhoppers, treehoppers, the larvae of sawflies, and the caterpillars
of many moths. Both the
Insect Table and the
Moth Table list many of
these species. Vertebrate animals also use
Rubus spp. as
sources of
food, particularly their fruits. The fruits of dewberries are sometimes
eaten by turtles, including
Clemmys
insculpta (Wood Turtle),
Terrapene
carolina (Eastern Box Turtle),
Terrapene ornata
(Ornate Box Turtle),
and
Kinosternum
subrubrum (Eastern Mud Turtle). Many mammals also feed
on the fruits of dewberries and other
Rubus spp.,
including the Black
Bear, Gray Fox, Red Fox, Raccoon, Opossum, Eastern Chipmunk, Red
Squirrel, Gray Squirrel, Fox Squirrel, White-Footed Mouse,
and
Woodland Deer Mouse. In wetland areas, muskrats feed on the roots to a
minor extent, while White-Tailed Deer and the Cottontail Rabbit feed on
the foliage and woody stems in a wider range of habitats. The fruits of
Rubus spp. are
also eaten by upland gamebirds and many songbirds (see
Bird Table). These various animals help to disperse seeds of the fruits
into new areas.
Photographic
Location: A sandy swamp at the Indiana Dunes State Park in
NW Indiana.
Comments: While most blackberries and raspberries produce
arching canes
up to 6' high, dewberries produce trailing stems that creep along the
ground. Compared to the upland species,
Rubus frondosus
(Common
Dewberry), Swamp Dewberry has leaflets with tips that are more blunt
and its stems are bristly, rather than prickly. The flowers of Swamp
Dewberry are also smaller in size (½-¾" in across) than those of Common
Dewberry (¾-1" across). Another species,
Rubus pubescens
(Dwarf
Raspberry), also produces trailing stems like a dewberry and it is
found in habitats that are similar to those of Swamp Raspberry. Dwarf
Raspberry usually has leaflets with more pointed tips than those of Swamp
Dewberry, and its stems have soft hairs, rather than sharp bristles or
prickles. Dwarf Raspberry usually produces its flowers individually,
rather than in small corymbs, and its mature fruits are red, rather
than dark purple or black. Another dewberry,
Rubus trivialis
(Southern
Dewberry), occurs in southern Illinois. This latter species differs
from Swamp Dewberry by having stems with both prickles and gland-tipped
hairs. Its non-flowering stems produce leaves with 5 leaflets, rather
than 3 leaflets, and its flowers are produced individually, rather than
in small corymbs.